After implantation, finger-like projections appear on the trophoblast called the chorionic villi which are surrounded by the uterine tissue and maternal blood.
The chorionic villi and uterine tissue become interdigitated with each other and jointly form a structural and functional unit between developing embryo (foetus) and maternal body called the placenta.
The placenta facilitates the supply of oxygen and nutrients to the embryo and also removal of carbon dioxide and excretory/waste materials produced by the embryo.
The placenta is connected to the embryo through an umbilical cord which helps in the transport of substances to and from the embryo.
Placenta also acts as an endocrine tissue and produces several hormones like human chorionic gonadotropin (hCG), human placental lactogen (hPL), estrogens, progestogens, etc.
In the later phase of pregnancy, a hormone called relaxin is also secreted by the ovary.
The hCG, hPL and relaxin are produced in women during pregnancy.
In addition, during pregnancy the levels of other hormones like estrogens, progestogens, cortisol, prolactin, thyroxine, etc. are increased several folds in the maternal blood.
Increased production of these hormones is essential for supporting the fetal growth, metabolic changes in the mother and maintenance of pregnancy.
Types of Mammalian Placenta
Types of placenta according to the nature of the foetal membranes taking part in the formation of the placenta:
Chorio-vitelline or “Yolk-sac” placenta-
It is a primitive type of placenta found in some of the marsupials. E.g., Opossum and Kangaroo.
In this type of placenta, the allantois remains comparatively tiny and never makes fusion with the chorion, while the yolk sac becomes very huge and combines broadly with the chorion.
Chorio-allantoic placenta-
In chorio-allantoic placenta, the yolk sac remains undeveloped.
The fusion found between the uterine wall and the embryo is lined by chorion and allantois.
Since the placenta is formed of chorion and allantois, it is termed as chorio-allantoic placenta. E.g., Parameles, Dasyurus.
Chorionic placenta-
The chorionic placenta is formed of a thickened layer of chorion containing sinuses filled with maternal blood.
Chorionic type of placenta is found in human beings.
The portion of the trophoblast which is nearer to the embryo is known as cytotrophoblast.
The more external lying part of the trophoblast is called syncytiotrophoblast as it is a syncytium of irregular strands with interstices in between.