The young pollen grain or male gametophyte has a centrally placed nucleus.
As it grows, the nucleus is pushed to one side and divided through an unequal mitotic division to form a small generative cell and a large vegetative cell or tube cell.
The generative cell is freely present in the cytoplasm of the vegetative cell.
Such a pollen grain is 2-celled and is shed like this in 60% of the angiosperms.
In the rest of the plants, the generative cell divides to form two non-motile male gametes.